Cars 2.Lightning McQueen, now a 4-time Piston Cup champion, returns home to Radiator Springs to enjoy some quiet time, but when Italian formula race car, Francesco Bernoulli, challenges McQueen to the newly created World Grand Prix, led by its creator Sir Miles Axlerod, he and his best friend Mater — along with Luigi, Guido, Fillmore, and Sarge — depart for Tokyo for the first race of the Grand Prix. Meanwhile, the lemons, who are led by weapons designer Professor Zündapp and an unknown mastermind, secretly plot to secure their oil profits by using an Electromagnetic pulse gun disguised as a camera (discovered by Finn on the oil platform) to trigger and destabilize the use of Allinol, a fuel that was created by Axlerod and required for racers to use in the Grand Prix. McMissile and his partner Holley Shiftwell attempt to meet with American spy car Rod "Torque" Redline at a World Grand Prix promotional event in Tokyo, to receive information about the mastermind. However, Redline is attacked by Zündapp's henchmen, but not before passing his information to Mater before he is captured, who is then mistaken to be the American contact of Holley and Finn. Before killing Redline, Professor Zündapp finds out that Mater was given the information.

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