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The series takes place in the fictitious city of Mariner Bay, California, which was built on an ancient demon burial ground. When the demons were accidentally released from their tomb in the desert, they threaten to destroy Mariner Bay. Therefore, a government organization called Lightspeed Rescue, headed by Captain William Mitchell, recruits four civilians and his own daughter to defend the city. Each of the four civilians chosen had a special area of expertise: Carter Grayson, the Red Lightspeed Ranger, was a fire fighter in the local fire department; Chad Lee, the Blue Lightspeed Ranger, worked as a lifeguard and marine animal trainer at a local aquarium and a marine amusement park; Joel Rawlings, the Green Lightspeed Ranger, was a stunt pilot; Kelsey Winslow, the Yellow Lightspeed Ranger, was an extreme sports athlete; and Dana Mitchell, the Captain’s daughter, who agreed to become the Pink Lightspeed Ranger, was a nurse and practiced medical arts. The five Power Rangers were aided by a team of scientists and engineers led by Miss Angela Fairweather, and operated out of the Lightspeed Aquabase, an underwater military compound that also deters the hydrophobic demons from directly attacking the base.

The five Rangers would be joined by Captain Mitchell’s (long thought to be dead) son, Ryan Mitchell, who would become the Titanium Ranger. Together, the six Rangers would prevail against the demon forces time after time, culminating in a final showdown where, in the Power Rangers tradition, all of the Zords and much of the weapons and other equipment that was used by the Power Rangers over the series was destroyed.

In addition, Lightspeed Rescue featured a reunion reappearance of characters from Power Rangers Lost Galaxy, when the villainess Trakeena comes to Earth to destroy it and the Lightspeed Rangers team up with the Galaxy Rangers.